Publications

Herraiz, J. L., Ferrón, H. G., Botella, H., Reolid, M., & Martínez-Pérez, C. (2026). The Iberian fossil record of† Otodus megalodon rejects Mediterranean dwarfism and supports nursery use. Biology Letters, 22(1): 20250640.    

Ferrón, H. G. (2025). QuAnTeTrack: Quantitative Analysis of Tetrapod Trackways. v0.1.0. GitHub.      

Sanchez-Sanchez, V., Sanisidro, O.,& Ferrón, H. G. (2025). Functional aspects of the headshield processes in ostracoderms. Palaeontology, 51(2), 310-322.     

Grohganz, M. Ballell, Ferrón, H. G., A., Rayfield, E.J., Johanson, Z, & Donoghue, P. C. (2024). Finite element and microstructural analyses indicate that pteraspid heterostracan oral plate microstructure was adapted to a mechanical function. Palaeontology, 67(6), e12733.     

Grohganz, M., Ferrón, H. G., Johanson, Z., & Donoghue, P. C. J. (2023). Testing hypotheses of pteraspid heterostracan feeding using computational fluid dynamics. Journal of Vertebrate Paleontology, 43(2), e2272974.    

Herraiz, J. L., Ferrón, H. G., Carbonell, Á., García-Sanz, I., Botella, H., & Martínez-Pérez,C. (2023). New data on the Late Miocene chondrichthyans from the Western Mediterranean region (Alcoy Basin, Eastern Spain). Journal of Vertebrate Paleontology, 43(4), e2307071.    

Gai, Z., Lin, X., Shan, X., Ferrón, H. G., & Donoghue, P. C. J. (2023). Postcranial disparity of galeaspids and the evolution of swimming speeds in stem-gnathostomes. National Science Review, nwad050.    

Ferrón, H. G. (2023). Illuminating the evolution of bioluminescence in sharks. Palaeontology, 66(1), e12641.    

Lovrenčić, L., Ferrón, H. G. , Grbin, D., & Maguire, I. (2022). Insight into the noble crayfish morphological diversity: a geometric morphometric approach. Knowledge & Management of Aquatic Ecosystems, (423), 9.      

Greif, M., Ferrón, H. G., & Klug, C. (2022). A new Meckel’s cartilage from the Devonian Hangenberg black shale in Morocco and its position in chondrichthyan jaw morphospace.  PeerJ ,  10 , e14418.      

Jobbins, M., Rücklin, M., Ferrón, H. G., & Klug, C. (2022). A new selenosteid placoderm from the Late Devonian of the eastern Anti-Atlas (Morocco) with preserved body outline and its ecomorphology. Frontiers in Ecology and Evolution, 10, 1007.      

Gai, Z., Li, Q., Ferrón, H. G., Keating, J. N., Wang, J., Donoghue, P. C., & Zhu, M. (2022). Galeaspid anatomy and the origin of vertebrate paired appendages. Nature, 609(7929), 959-963.      

Ferrón, H. G., & Donoghue, P. C. (2022). Evolutionary analysis of swimming speed in early vertebrates challenges the ‘New Head Hypothesis’. Communications Biology, 5(1), 863.      

Ferrón, H. G., Ballell, A., Botella, H., & Martinez-Perez, C. (2022). Biomechanics of Machaeracanthus pectoral fin spines provide evidence for distinctive spine function and lifestyle among early chondrichthyans. Journal of Vertebrate Paleontology, 41(6), e2090260.       

Herraiz, J. L., Carrillo-Briceño, J. D., Ferrón, H. G., Adnet, S., Botella, H., & Martínez-Pérez, C. (2022). First fossil record (Middle Miocene) of the viper shark Trigonognathus Mochizuki and Ohe, 1990, in the Mediterranean realm. Journal of Vertebrate Paleontology, 42(1), e2114360.      

Ferrón, H. G., & Palacios-Abella, J. F. (2022). Grouping behaviour impacts on the parasitic pressure and squamation of sharks. Proceedings of the Royal Society B, 289(1975), 20220093.    

Figueirido, B., Serrano, F. J., Pérez-Ramos, A., Esteban, J. M., Ferrón, H. G., & Martín-Serra, A. (2022). Body-axis organization in tetrapods: a model-system to disentangle the developmental origins of convergent evolution in deep time. Biology Letters, 18(4), 20220047.    

Ferrón, H. G., Martínez-Pérez, C., Rahman, I. A., Selles de Lucas, V., Botella, H., & Donoghue, P. C. (2021). Functional assessment of morphological homoplasy in stem-gnathostomes. Proceedings of the Royal Society B, 288(1943), 20202719.    

Ballell, A., & Ferrón, H. G. (2021). Biomechanical insights into the dentition of megatooth sharks (Lamniformes: Otodontidae). Scientific Reports, 11(1), 1232.    

Herraiz, J. L., Ribé, J., Botella, H., Martínez-Pérez, C., & Ferrón, H. G. (2020). Use of nursery areas by the extinct megatooth shark Otodus megalodon (Chondrichthyes: Lamniformes). Biology Letters, 16(11), 20200746.   

Ferrón, H. G., Martínez-Pérez, C., Rahman, I. A., de Lucas, V. S., Botella, H., & Donoghue, P. C. (2020). Computational fluid dynamics suggests ecological diversification among stem-gnathostomes. Current Biology, 30(23), 4808-4813.   

Cooper, J. A., Pimiento, C., Ferrón, H. G., & Benton, M. J. (2020). Body dimensions of the extinct giant shark Otodus megalodon: a 2D reconstruction. Scientific reports, 10(1), 14596.    

Ferrón, H. G., Greenwood, J. M., Deline, B., Martínez‐Pérez, C., Botella, H., Sansom, R. S., Ruta, M., & Donoghue, P. C. (2020). Categorical versus geometric morphometric approaches to characterizing the evolution of morphological disparity in Osteostraci (Vertebrata, stem Gnathostomata). Palaeontology, 63(5), 717-732.   

Ferrón, H. G. (2019). Evidence of endothermy in the extinct macropredatory osteichthyan Xiphactinus audax (Teleostei, Ichthyodectiformes). Journal of Vertebrate Paleontology, 39(6), e1724123.   

Manzanares, E., Escudero-Mozo, M. J., Ferrón, H. G., Martínez-Pérez, C., & Botella, H. (2020). Middle Triassic sharks from the Catalan Coastal ranges (NE Spain) and faunal colonization patterns during the westward transgression of Tethys. Palaeogeography, Palaeoclimatology, Palaeoecology, 539, 109489.    

Martínez-Pérez, C., Navas-Parejo, P., Aaron Lara-Pena, R., Ferrón, H. G., Palafox-Reyes, J., & Botella, H. (2019). Late Devonian (Famennian) Chondrichthyes from Mexico. Journal of Vertebrate Paleontology, 39(6), e1764008.   

Ferrón, H. G., Herraiz, J. L., Botella, H., & Martínez-Pérez, C. (2021). Pre-Messinian ecological diversity of Mediterranean sharks revealed by the study of their dermal denticles. Spanish Journal of Palaeontology, 34(2), 289-298.    

Martinez-Perez, C., Martin-Lazaro, A., Ferrón, H. G., Kirstein, M., Donoghue, P. C., & Botella, H. (2018). Vascular structure of the earliest shark teeth. Acta Geologica Polonica, 68(3), 457-465.   

Ferrón, H. G., Holgado, B., Liston, J. J., Martínez‐Pérez, C., & Botella, H. (2018). Assessing metabolic constraints on the maximum body size of actinopterygians: locomotion energetics of Leedsichthys problematicus (Actinopterygii, Pachycormiformes). Palaeontology, 61(5), 775-783.   

Manzanares, E., Pla, C., Ferrón, H. G., & Botella, H. (2018). Middle-Late Triassic chondrichthyans remains from the Betic Range (Spain). Journal of Iberian Geology, 44, 129-138.   

Ferrón, H. G., Martínez‐Pérez, C., Turner, S., Manzanares, E., & Botella, H. (2018). Patterns of ecological diversification in thelodonts. Palaeontology, 61(2), 303-315.   

Ferrón, H. G., Paredes-Aliaga, M. V., Martínez-Pérez, C., & Botella, H. (2018). Bioluminescent-like squamation in the galeomorph shark Apristurus ampliceps (Chondrichthyes: Elasmobranchii). Contributions to Zoology, 87(3), 187-196.    

Ferrón, H. G., & Martinez-Perez, C. (2017). Life in the Palaeozoic: an overview of land and sea ecosystems. Bulletin of Geosciences, 92(4), 439-442.    

Ferrón, H. G., Martínez-Pérez, C., & Botella, H. (2017). Ecomorphological inferences in early vertebrates: reconstructing Dunkleosteus terrelli (Arthrodira, Placodermi) caudal fin from palaeoecological data. PeerJ, 5, e4081.    

Ferrón, H. G. (2017). Regional endothermy as a trigger for gigantism in some extinct macropredatory sharks. PloS one, 12(9), e0185185.    

Martínez-Pérez, C., Carrillo-Briceño, J. D., Esparza, C., Ferrón, H. G., Manzanares, E., Hammann, C., & Botella, H. (2018). A Serravallian (Middle Miocene) shark fauna from Southeastern Spain and its palaeoenvironment significance. Historical Biology, 30(3), 422-432.   

Ferrón, H. G., Martínez-Pérez, C., & Botella, H. (2018). The evolution of gigantism in active marine predators. Historical Biology, 30(5), 712-716.   

Ferrón, H. G., & Botella, H. (2017). Squamation and ecology of thelodonts. PloS one, 12(2), e0172781.    

Manzanares, E., Pla, C., Martínez-Pérez, C., Ferrón, H. G., & Botella, H. (2017). Lonchidion derenzii, sp. nov., a new lonchidiid shark (Chondrichthyes, Hybodontiforms) from the Upper Triassic of Spain, with remarks on lonchidiid enameloid. Journal of Vertebrate Paleontology, 37(1), e1253585.   

Ferrón, H. G., Pla, C., Martínez-Pérez, C., Escudero-Mozo, M. J., & Botella, H. (2014). Morphometric Discriminant Analysis of isolated chondrichthyan scales for palaeoecological inferences: the Middle Triassic of the Iberian Chain (Spain) as a case of study. Journal of Iberian Geology, 40, 87-97.   

Botella, H., Manzanares, E., Ferrón, H. G., & Martínez-Pérez, C. (2014). Obruchevacanthus ireneae gen. et sp. nov., a new ischnacanthiform (Acanthodii) from the Lower Devonian of Spain. Paleontological Journal, 48, 1067-1076.